In the aftermath of the Black Cross Army's surprise attack on EAGLE bases throughout Japan, five surviving agents band together to face them as the Gorenger.
A band of space pirates known as the Gokaiger arrive on Earth in pursuit of a legendary treasure tied to the 34 previous Sentai teams of the planet, who had all sacrificed their powers to stop the Zangyack Empire.
The Gokaigers receive a clue to the greatest treasure in that they should search for someone wearing black clothing. When they come across a student in a black school uniform, however, the boy steals one of Captain Marvelous' Ranger Keys.
After the Gokai Galleon makes a crash landing in a volcano, the Gokaigers are attacked by Action Commander Salamandam. Elsewhere, Captain Marvelous and Don Dogoier meet the prophesied man dressed in black.
Joe's past is revealed when Action Commander Zodomas is sent to fight the Gokaigers.
When Navi reveals that the next clue to the Greatest Treasure in the Universe is in the hands of the police, Captain Marvelous seeks out a police station only to be arrested by the Dekarangers.
Luka goes shopping for jewelry until she comes across a young woman whose father is very wealthy, leading Luka to want to go on a heist while undercover as a maid.
With the next clue from Navi saying that they need to find the tiger child, the Gokaigers face off against Pachacamac XIII. When he takes away their weapons, Doc and Ahim find their hand-to-hand skills are lacking and start training under Jan Kandou, the former GekiRed.
The Zangyack Empire sends the Sneak Brothers to spy on the Gokaigers.
In a race to find the floating island before the Zangyack forces do, the Gokai Galleon crew instead finds a veterinarian.
Joe and Luka infiltrate the Green Fleet, the Zangyack Empire's special forces, to learn more about their Gigarollium Cannon.
With Navi's next clue being "Sa-mu-r-aa-i", the Gokai Galleon crew seeks out a kendo club until they are approached by a girl who claims to be the samurai known as Shinken Red. Joe begins training with her to face off against his old comrade Barizorg.
Joe goes off to face Barizorg alone in an attempt to make him remember his past as Kaoru assists the Gokaigers in their fight against the Zangyack Empire.
88 constellations that exist in the universe have fallen into the hands of the evil Space Shogunate Jark Matter. However, the insurrection army Rebellion and three Kyurangers chosen by the Kyutama are opposing them to protect the people. By an accident, an adventurer traveling through space, Lucky witnesses Chameleon Green, Oushi Black, and Kajiki Yellow fighting on the planet Crotus. Lucky decides to become the 4th Kyuranger and they set out to planet Jagjag to search for the Kyutama.
The search is on for the remaining four Kyurangers. Lucky lands on Planet Jigama, and he encounters Balance and Nāga as they lead around Jark Matter. These two people call themselves the "Phantom Thief Group BN", stealing only from Jark Matter. Lucky hears their next aim is to steal a jewel from Jark Matter's space ship Moraimarz, acting to test their luck together. However, they meet the first Daikaan commander, Gamettsui, who Balance betrayed. Elsewhere, Jark Matter's general, Eriedrone, appears before Hammy and the others.
On the desert planet of Needle, a danger of explosion has been identified due to Jark Matter sucking up the planet's "Planesium" energy. To prevent the explosion, the Spaceship Moraimarz that's sucking up the Planesium must be destroyed. Lucky and the others go down to Planet Needle to fight against Jark Matter. There, a man with a Kyutama named Stinger appears, and transforms into Sasori Orange. Champ sees Stinger's appearance, says something about "avenging the Doctor" and rushes in, shaking off the other Kyurangers trying to restrain him. It seems Stinger and Champ have a past together.
The Orion comes to Earth. There's always one Daikaan on every planet, but there are multiple Daikaans confirmed to be on board the Moraimarz on Earth. Lucky and the others head down to Earth, where the Daikaan known as Yumepakkun steals and eats the dreams of children. Lucky and the others transform and fight as the Kyurangers. Raptor dreams of fighting together with the Kyurangers, and her wild ideas and fascinations with fighting are only getting bigger. Yumepakkun invades Raptor's big dreams and destroys them with fears. Spada and the others believe Raptor's dreams are dangerous and oppose them, but Lucky thinks otherwise.
Once upon a time, there were five great nations, Shugoddam, N'Kosopa, Ishabana, Gokkan, and Toufu. Each country had its own ruler, who was allowed to fight by arming oneself in the form of a warrior with one's sword. According to tradition, it was said that the Earth Empire Bugnarok, an enemy supposed to have been defeated once 2,000 years ago, would rise again this year, and they were to fight against it. The rulers of the five countries had gathered in Shugoddam, the most powerful country in the world, to form an alliance in preparation for the return of Bugnarok...
The country "Shugoddam" where the guardian deity resides is in a pinch by the "Bugnarak" army revived from the emperor, but it is saved by Gira, who transforms into Kuwagata Ohger. However, he is chased by Shugoddam's soldiers for defying King Rcules. The one who helped Gira was the king of N'Kosopa, Yanma.
With the help of the people of N'Kosopa, Yanma and Gira defeat the enormous Bodarujim. The one who suddenly kidnapped them was Hymeno Ran, the queen of "Ishabana". Hymeno, who has a super selfish personality and wields Gira and Yanma, actually has a secret...?
Gira and Yanma learn of Hymeno's unusual selfishness and hidden considerations for the nation. In front of the three rulers, one of the three major guardian deities "God Kabuto" appears. The one who was riding God Kabuto was the lord of "Toufu", Kaguragi Dybowski. What is the nature of Kaguragi, who is thinking about something bad...?